Reginaldus Le Fowler, of Shryvenham, Co. Berks, is the first one of the line that can be definitely located. He d. 29 Edward I, 1301. In the Calendaium Genealogicum Henry III and Edward I., Master of the Rolls, Vol. 2, p. 602, we find the following: "Reginaldus Le Fowler de Shryvenham. Inquisition Post mortem: Johanes Le Fowler de Shryvenham filius praedicti Reginaldi est propinquior haeres ejus et est aetatis triginta duorum annorum amplius, 29 Edward I, 1301. Berks."
Master of the Rolls, Vol. 2, p. 602...
"According to certain accounts, the family of Fowler is of Norman descent and went into England with William the Conqueror in 1066. The earliest record of the name in England is that of Richard Fowler of Foxley, County of Buckingham, who accompanied Richard Coeur de Lion (Richard the Lionhearted) to the Holy Land in 1191. For his service he won the crest used with the family coat of arms and a a grant of lands at Abbey Cwm-Hir, County of Radnor, Wales. From Richard Fowler were descended the families of that name which were to be found at early dates in England, in the counties of Buckingham, Lancaster, Wilts, Stafford, Oxford, Gloucester, and Shrops, and also in Wales and Ireland. They appear to have been, for the most part, of the landed gentry and yeomanry of the British Isles."